general info about Theriologia Ukrainica

Novitates Theriologicae

ISSN 2709-491X (print) • ISSN 2709-4928 (online)

2026 • Vol. 18 • Contents of volume >>>


 

bibliography

download pdfLobkov, V., Y. Oleinik. 2026. A brief history and directions of theriological research in the North-Western Black Sea Region. Novitates Theriologicae, 18: 320–333. https://doi.org/10.53452/nt1828

   

title

A brief history and directions of theriological research in the North-Western Black Sea Region

author(s)

Vladimir Lobkov (I. I. Mechnikov Odesa National University, Odesa, Ukraine)
https://orcid.org/0000-0003-4707-9871
Yurii Oleinik (I. I. Mechnikov Odesa National University, Odesa, Ukraine)
https://orcid.org/0000-0002-0033-0718

abstract

The history of the study of mammals in the Black Sea Region covers about 180 years, starting with the work of Professor Nordmann. Since the end of the 19th century, theriological research in the region had expanded significantly, covering almost all sections of modern theriology by the end of the last century. As a result of long-term and systematic studies of the second half of the 20th century and at the beginning of the 21st century, theriologists (Gursky, Lobkov, and others) achieved the greatest success in the field of population ecology and morphology of some representatives of the Rodentia and Carnivora orders. Within the framework of these works, V. Lobkov developed the concept of dynamics of mammalian numbers with the participation of intrapopulation mechanisms of birth control, which is based on the connection of disintegration of the spatial and ethological structure of the population with the formation of outbred generations. On the basis of morphological studies, methods for determining the age of a number of predatory mammals and of rodents have been developed and verified. The essay briefly highlights the contribution of mammal experts of Odesa University and other institutions to the formation of scientific directions in the study of animals of the North-Western Black Sea Region. At the end, there is an annotated list of about 100 scientific papers that reflect the main stages and achievements in the study of the modern mammal fauna in the region.

language

Ukrainian, with English summary

   

references

Andrievskii, O. M., Y. M. Oleinik, A. S. Asmanska. 2012. Molecular forms of carboxyl esterases of the organs and tissue of ground squirrel Spermophilus suslicus (Guld.) in a late postnatal period of ontogenesis. Odesa National University Herald. Series Biology, 17 (3): 7–18. [Russian] https://doi.org/10.18524/2077-1746.2013.2(31).44603
Andrievskii, O. M., Y. M. Oleinik. 2013. Molecular forms of carboxylesterases of the organs and tissue of ground squirrel Spermophilus suslicus (Guld.) in the embryonic and а early postnatal period. Odesa National University Herald. Biology, 18 (2): 25–37. [Russian]
Archipov, A. 2012. Records of Some Mammals Included in Red Data Book of Ukraine in the Vicinity of Kuchurhansky Lyman of Odesa Region. Vestnik Zoologii, 46 (4): 336. [Russian]
Barkaszi, Z., O. Gaydash. 2019. Zoological collections: amassment, inventory, identification (report on the XXIV Theriological School). Theriologia Ukrainica, 17: 119–125. https://doi.org/10.15407/pts2019.17.119
Brauner, A. A. 2005. Seasonal migrations of wild ungulates. Bulletin of the A. A. Brauner's Museum Fund, 2 (3): 4–7. [Russian] URL
Brauner, A. A. 1923. Agricultural Zoology. State Publishing House of Ukraine, Odesa,1–413. [Russian].
Chernyaev, N. 1857. Description of ground squirrels living in southern Russia and methods of their extermination. St. Petersburg, 1–54. [Russian]
Dyatlova, O., M. Rozhenko. 2017. Records and peculiarities of distribution of the European mole (Talpa europaea) in the Lower Dniester area. Proceedings of the Theriological School,15: 94–96. [Ukrainian] https://doi.org/10.15407/ptt2017.15.094
Fedorchenko, A. A., I. V. Zagorodniuk. 1994. Mice of the genus Sylvaemus from Lower Danube Region. Communication 2. Distribution and abundance. Vestnik zoologii, 28 (4–5): 55–64. [Russian] URL
Fedorchenko, A., V. Tkach. 1998a. Bats of the Danube Delta. Proceedings of the Theriological School. European Bat Night '98 in Ukraine. 1: 87–89. [Russian]  URL
Godlevskaya, E., P. Panchenko, O. Formaniuk. 2008. New data on cave-dwelling bats of Odesa city vicinities. Proceedings of the Theriological School, 9: 93–101. [Russian]
Godlevska, L., P. Panchenko, S. Rebrov, [et al.]. 2018. Findings of bats in the territory of the Black Sea region of Ukraine (AR Crimea, Mykolaiv, Odesa, Kherson regions). Materials for the 4th edition of the Red Book of Ukraine. Animal World. Kyiv, 196–203. [Ukrainian]
Gursky, I. G. 1968. Wolf in the steppe zone of Ukraine and its impact on wild and domestic animals. Studying the Nature of the Steppes. Materials of Symposium. Odesa, 158–161. [Russian]
Gursky, I. G. 1970. Bite shape and abrasion of teeth. Vestnik zoologii, № 5: 56–60. [Russian]
Gursky, I. G. 1972. Distribution of animals in the northwestern Black Sea region and in the Crimea in the 17th–19th centuries. In: Kirikov, S. V. (ed.). Changes in the Population and Habitat of Animals in the European Part of the USSR in the 16th–19th Centuries. Conference proceedings, December 1972 MOIP, Moscow, 19–22. [Russian]
Gursky, I. G. 1973. Determination of the age of the wolf (Canis lupus L.) by the skull. Vestnik zoologii, No. 3: 1–69. [Russian]
Gursky, I. G. 1975. Wolf-dog hybridization in nature. Bulletin MOIP. Dept. biological, No. 1: 131–136. [Russian]
Gursky, I. G., V. A. Urban. 1976. Fox and other predatory animals in Odesa. Bulletin MOIP. Dept. biological, No. 5: 107–108. [Russian]
Gursky, I. G. 1978. Wolf in the northwestern Black Sea region (habitat, population structure, reproduction). Bulletin MOIP. Dept. biological, No. 3: 29–38. [Russian]
Gursky, I. G. 1982. Wolf in Ukraine and Moldova. Hunting and game economy, No. 3: 10–11. [Russian]
Gursky, I. G. 1985. The number and characteristics of the way of life by region. Ukraine and Moldova. In: Bibikov, I. D. (ed.). Wolf. Nauka, Moscow, 487–493. [Russian]
Gursky, I. G. 1989. New data on the number of wolves and its regulation in Ukraine. In: Ecology, behavior and Management of Wolf Populations. Moscow, 55–57. [Russian]
Kivganov, D. A., I. Y. Omelchuk, Y. N. Oleinik. 2004. About the discovery of the Mediterranean bat, Pipistrellus kuhlii, on island Zmeinyy. Vestnik zoology, 38 (4): 94. [Russian]
Klevezal, G. A., V. A. Lobkov, D. V. Shchepotkin. 2021. Hibernation records on the surface of rodent incisors: intraspecific variations and interspecific differences. Russian Journal of Zoology, 100 (5): 524–539. [Russian]
List of scientific publications of Gurski I. G. 2008. Bulletin of the A. A. Brauner's Museum Fund, 5 (2): 9–11. URL
Lobkov, V. A. 1977. On seasonal changes in the activity of the speckled ground squirrel (Citellus suslicus). Russian Journal of Zoology, 56 (12): 1900–1904. [Russian]
Lobkov, V. A. 1978. On seasonal changes in the activity of the speckled ground squirrel (Citellus suslicus). Russian Journal of Zoology, 57(12): 1897–1899. [Russian]
Lobkov, V. A. 1983. Reproduction of the spotted ground squirrel (Citellus suslicus Guld.) in the anthropogenic landscape of the northwestern Black Sea region. Ecology, (2): 44–49. [Russian]
Lobkov, V. A. 1984. Group labeling of young speckled ground squirrels with tetracycline to study their dispersal. Russian Journal of Zoology, 63 (2): 309–311. [Russian]
Lobkov, V. A. 1991. Changes in the number and sex structure of the settlements of the speckled ground squirrel (Citellus suslicus) in the spring. Russian Journal of Zoology, 70 (10): 114–122. [Russian]
Lobkov, V. A. 1993. Dynamics of settlements of the spotted ground squirrel (Citellus suslicus) in the Northwestern Black Sea region. Mammals of Ukraine, 105–113. [Russian]
Lobkov, V. A., N. V. Rozhenko. 1998. Findings of the pine marten (Martes martes L.) in the steppe zone in southwestern Ukraine. In: Studies on the Diversity of the Animal Word: Proceedings of the OSU Zoological Museum, 3: 188–189. [Russian]
Lobkov, V. A. 1999. Spotted ground squirrel of the North-Western Black Sea region: biology, functioning of populations. Astroprint, Odesa, 1–270. [Russian]
Lobkov, V. A. 2003. Population mechanisms of adaptation of the speckled ground squirrel Spermophilus suslicus (Mammalia, Rodentia) to living in agrocenoses. Bulletin of Dnipropetrovsk University. Biology. Ecology, 1 (11): 178–183. [Russian]
Lobkov, V. A. 2006a. Mortality in populations of the speckled ground squirrel (Spermorhilus suslicus Rodentia, Sciuridae) of the Northwestern Black Sea region. Russian Journal of Zoology, 85 (10): 1247–1256. [Russian]
Lobkov, V. A. 2006b. Ecological causes of changes in the abundance and distribution of the speckled ground squirrel Spermorilus suslicus (Guldenstaedt, 1770). Bulletin of MOIP, Dept. biological, 111 (5): 59–64. [Russian]
Lobkov, V. A. 2009. On the similarity and nature of phenomena observed during the formation of spatial groups of speckled ground squirrel (Spermophilus suslicus Guld.) and muskrat (Ondatra zibethicus L.). Bulletin of the Odesa National University. Series Biology, 14 (14): 78–87. [Russian]
Lobkov, V. A. 2010. Regularities of the existence of isolated settlements of the speckled ground squirrel (Spermophilus suslicus, Rodentia, Sciuridae) in the North-Western Black Sea region. 1. Spatial distribution of individuals. Russian Journal of Zoology, 89 (11): 1374–1379. [Russian]
Lobkov, V. A. 2011. Regularities of the existence of isolated settlements of the speckled ground squirrel (Spermophilus suslicus, Rodentia, Sciuridae) in the North-Western Black Sea region. 2. Intrapopulation regulation of the number. Zoologicheskii zhurnal, 90 (3): 342–350. [Russian]
Lobkov, V. A. 2013. Dynamics of foxes reproduction (Vulpes vulpes Linnaeus, 1758) in conditions of decreasing anthropogenous influence. Odesa National University Herald. Biology, 18 (1): 67–71. [Russian] https://doi.org/10.18524/2077-1746.2013.1(30).44576
Lobkov, V. A. 2016. Intrapopulation regulation of the number of mammals. Odesa National University, Odesa, 1–237. [Russian]
Lobkov, V. A. 2019. On the results and prospects of nature protection and the conservation of rare species of animals and plants. News of the Askania-Nova Biosphere Reserve, 21: 73–79. [Russian]
Lobkov, V. A. 2020. Some reflections on the current and conservation status of the wildcat (Felis sylvestris) in the northwestern Black Sea region. Bulletin of the A. A. Brauner's Museum Fund, 17 (2): 31–33. [Russian] URL
Lobkov, V. A. 2023a. The reduction of the size of wolves of the same age of the northwestern black sea region in the 21st century. Russian Journal of Ecology, 54 (1): 70–76. [Russian] https://doi.org/10.1134/S1067413623010071
Lobkov, V. A. 2023b. On the patterns and causes of chronographic variability of morphological characteristics in some mammals: The orders Rodentia and Carnivora. Russian Journal of Zoology, 102 (10): 1172–1188. [Russian] https://doi.org/10.31857/S0044513423100069
Lobkov, V. A., D. V. Schepotkin. 2024. Recording the peculiarities of life activities fixed on the surface of the incisors of the spotted ground squirrel (Spermophilus suslicus, Rodentia, Sciuridae). Russian Journal of Zoology, 103 (6): 46–59. [Russian] https://doi.org/10.31857/S0044513424060032
Lyashevskaya, T. V. 1992. Some data on the fauna and ecology of bats in the Odesa region. Scientific Works of the Zoological Museum of Odesa State University I. I. Mechnikov. Animal Fauna and Ecology, 1: 38–39. [Russian]
Makhno, A. 1889. Spotted ground squirrel (Spermophilus guttatus). Digest of Kherson Zemstvo, (4): 68–73. [Russian]
Oleinik, Y. N., V. A. Lobkov. 1990. Changes in the state of the thyroid gland of the spotted gopher in different settlements of the northwestern Black Sea region. Spotted Suslik in the North-Western Black Sea Region. Institute of Zoology AS of Ukr. SSR, Preprint 90.17: 3–6. [Russian]
Oleinik, Y. N. 1999. Structural organization and morphometric characteristics of the thyroid gland of the spotted gopher (Spermophillus suslicus). Development of Zoological Research at Odesa University. Acad. D.K. Tretyakov and His Scientific School, 244–249. [Russian]
Oleinik, Y. N. 2003a. The structural organization and development of the thyroid gland of the spotted ground squirrel (Spermophillus suslicus Guld.) in the early postnatal period. Odesa National University Herald. Series biology, 8(1): 139–144. [Russian]
Oleinik, Y. N. 2003b. Morpho-functional state of the thyroid gland of the spotted gopher (Spermophilus suslicus Guldenstaedt, 1770) in settlements of different densities. Herald of Dnipropetrovsk University. Series Biology. Ecology, 1 (11): 192–197. [Russian]
Oleinik, Y. N., V. A. Lobkov. 2003. Postnatal development of speckled ground squirrel (Spermophillus suslicus Guld.). Odesa National University Herald.Series Biology, 8 (6): 131–137. [Russian]
Oleinik, Y. N. 2004. The structural organization and seasonal changes of the thyroid gland of the male spotted ground squirrel (Spermophillus suslicus Guld.). Odesa National University Herald. Series biology, 9 (1): 185–192. [Russian]
Oleynik, Y. N., N. V. Rozhenko. 2011. Essay of theriofauna of the delta area of river Dnister. Bulletin of the A. A. Brauner's Museum Fund, 8(4): 1–28. [Russian] URL
Oleinik, Y. N. 2020. About the encounters of the wildcat (Felis silvestris Schreber, 1777) in the south-west of Ukraine (Odesa region) at the beginning of the XXI century. Bulletin of the A. A. Brauner's Museum Fund, 17 (2): 11–30. [Russian] URL
Oleinik, Y. N. 2021. Sex-age composition of the common fox (Vulpes vulpes L.) in the Odesa region. Bulletin of the A. A. Brauner's Museum Fund, 18 (1): 31–35. [Russian] URL
Oleinik, Y. 2024. Age determination of the wildcat (Felis silvestris): a case study of a sample from the north-western black sea region (Ukraine). Theriologia Ukrainica, 28: 69–83. https://doi.org/10.53452/TU2806
Panchenko, P., L. Godlevska. 2018. Data on chiropterofauna of the Northern Black Sea region and results of the work of bat contact-centres. Theriologia Ukrainica, 16: 120–126. [Ukrainian] https://doi.org/10.15407/pts2018.16.120
Potapov, O. 2000. New records of the European wild cat (Felis silvestris) in the Danube delta. Vestnik Zoologii, 34 (6): 88. [Russian]
Puzanov, I. I. 1956. Patterns of distribution of systematic features of the southern populations of small and speckled ground squirrels. Scientific Yearbook of Odssa University, Odesa, 257–258. [Russian]
Puzanov, I. I. 1958. On some regularities in the distribution of systematic features of the speckled ground squirrel. Problems of Land Zoogeography. Publishing House of Lviv University, Lviv, 203–209. [Russian]
Ramenskiy, S. Y., V. S. Smirnov, I. G. Gursky, M. P. Pavlov 1985. The increase in the size of wolves of the same age in the XX century. In: Bibikov, I. D. (ed.). Wolf. Nauka, Moscow, 261–266. [Russian]
Rozhenko, N. V., A. M. Volokh. 1998. Settlement of the raccoon dog in the Northern Azov and Black Sea regions.In: Studies on the Diversity of the Animal Word: Proceedings of the OSU Zoological Museum, 3: 133–137. [Russian]
Rozhenko, N. V. 2000. On recent findings of the wildcat (Felis silvestris Schr.) in the lower reaches of the Dnister River. Vestnik Zoologii, 34 (6): 80. [Ukrainian]
Rozhenko, N. V., A. M. Volokh. 2000. Appearance the golden jackal (Canis aureus) in the South of Ukraine. Vestnik zoologii, 34 (1–2): 125–129. [Ukrainian]
Rozhenko, N. V. 2002. Some results of research on the ecology of the red fox in the Black Sea region. Visnyk Lviv University, 30: 84–89. [Ukrainian]
Rozhenko, N. V. 2004. Dynamics and current state of the fauna of predatory mammals in the Dniester and Danube deltas.Scientific Notes of the Tavrichesky National University. Series Biology, Chemistry, 17 (56). No. 2: 115–120. [Ukrainian]
Rozhenko, N. 2006. Feeding of some carnivorous mammals in anthropogenic landscape of the Black Sea region. Proceedings of Theriological School, 8: 191–200. [Ukrainian]
Rozhenko, N. V. 2007. Dynamics of the abundance of the raccoon dog (Nyctereutes procyonoydes Matsch.) in the Ukrainian Black Sea region. Herald of hunting, 4 (1): 50. [Russian]
Rozhenko, N. V. 2010. First Record of Golden Jackal,Canis aureus(Carnivora, Canidae), in the Lower Danube. Vestnik Zoologii, 44 (3): 208 [Ukrainian]
Rozenko, N., A. Volokh 2010. The golden jackal (Canis aureus L., 1758) as a new species in the fauna of Ukraine. Beiträge zur Jagd- & Wildforschung, 34: 237–246.
Rozhenko, M. V. 2011. State of the wildcat population in the North-Western Black Sea region. Ecological Herald, No. 4 (68): 6. [Ukrainian]
Rozhenko, M. 2015. Population Dynamics and Distribution of the European Mink (Mustela lutreola) in the Lower Dnister Region in 2009–2014. Proceedings of the Theriological School, 13: 35–36. [Ukrainian] https://doi.org/10.15407/ptt2015.13.035
Rozhenko, N. 2021. The current state of the golden jackal population in the Lower Dnister National Nature Park. Theriologia Ukrainica, 21: 109–113 https://doi.org/10.15407/TU2109
Rusev, I. 1998. Mammal fauna and the history of research in the Dniester floodplains protected area. Nature Protection in Ukraine, 4 (1): 58–61. [Russian]
Rusev, I. 2020. The golden jackal (Canis aureus) in the Tuzlivski Lymany National Nature Park. Theriologia Ukrainica, 20: 46–57. [Ukrainian] https://doi.org/10.15407/TU2007
Rusev, I. 2026. The role of the Odesa Anti-Plague Station in the system for monitoring natural foci of zoonoses (a historical overview). Novitates Theriologicae, 18: 150–159. [Ukrainian] https://doi.org/10.53452/nt1809
Stronen, A. V., B. Jedrzejewska, C. Pertoldi, [et al.]. 2013. North-south differentiation and a region of high diversity in European wolves (Canis lupus). PLOS One, 8 (10): e76454. https://doi.org/10.1371/journal.pone.0076454
Tkach, V., A. Fedorchenko. 1998b. Records of Pipistrellus kuhlion the South of Ukraine. In: Zagorodniuk, I. (ed.). European Bat Night '98 in Ukraine, Kyiv, 150–152. (Series: Proceedings of the Theriological School; Vol. 1). [Russian] URL
Tokar, G. A., A. S. Matveev. 2010. Findings of a wildcat in the Ukrainian part of the Danube Delta in 2006–2007. Nature Conservation in Ukraine, 16 (1): 50–52. [Russian]
Trunova, Y. E., V. A. Lobkov. 1997. Recording the hibernation peculiarities by incisor dentin in spotted suslik Spermophilus suslicus (Sciuridae, Rodentia). Russian Journal of Zoology, 1 (3): 352–359. [Russian] https://doi.org/10.4098/AT.arch.99-14
Trunova, Y. E., V. A. Lobkov, G. A. Klevezal. 1999. The record of the reproductive cycle in the incisor dentine of spotted souslik Spermophilus suslicus. Acta Theriologica, 44 (2): 161–171.
Volokh, A. M., N. V. Rozhenko, V. A. Lobkov. 1998. The first meeting of the common jackal (Canis aureus L.) in the south-west of Ukraine. Scientific Works of the Zoological Museum of the Odesa State University, 5: 187–188. [Russian].
Volokh, A. M., N. V. Rozhenko. 2004. Biotopic distribution and shelters of the raccoon dog (Nyctereutes procyonoides Matsch.) in southern Ukraine. Russian Journal of Zoology, 83 (5): 635–638. [Russian].
Volyansky, Y. E. 1967a. Wintering of bats in the vicinity of Odesa. Vestnik zoologii, 1 (1): 77–78. [Russian]
Volyansky, Y. E. 1967b. Beech marten (Martes foina Erxl.) in Odesa. Vestnik zoologii, 1 (6): 84–86. [Russian]
Volokh, A., N. Rozenko. 2011. Dynamics of the range and population of the European mink (Mustela lutreolaL., 1761) in Ukraine. Beiträge zur Jagd- und Wildforschung, 36: 425–435.
Volokh, A., N. Rozenko. 2016. Modern Distribution and Morphology of the Golden Jackal (Canis aureus) in Ukraine. Beiträge zur Jagd- und Wildforschung, 41: 307–318.
Yurchenko, O., D. Dubina, D. Sokolovskyi, O. Gaidash. 2017. Possible zoonotic viral threats associated with bats in southern Ukraine. Proceedings of the Theriological School, 15: 150–153. https://doi.org/10.15407/ptt2017.15.150
Zagorodniuk, I. V., A. A. Fedorchenko 1993. Mice of the genus Sylvaemus in the Lower Danube Region. Communication 1. Taxonomy and diagnostics. Vestnik zoologii, 27 (3): 41–49. [Russian] URL

date

online: 30.01.2026; print: 30.01.2026


 


to main page of journal >>>

created: 25.01.2026
updated: 18.04.2026

Locations of visitors to this page