Theriologia Ukrainica
(former Proceedings of the Theriological School)

ISSN 2616-7379 (print) ISSN 2617-1120 (online)

2019 Vol. 17 Contents of volume >>>

download pdfStetsula, N. The dynamic component in the mammal fauna of the Skole Beskyds (Ukrainian Carpathians). Theriologia Ukrainica. 2019. Vol. 17: 43-54.



The dynamic component in the mammal fauna of the Skole Beskyds (Ukrainian Carpathians)


Stetsula, N.


Drohobych Ivan Franko State Pedagogical University (Drohobych, Ukraine)


Theriologia Ukrainica. 2019. Vol. 17: 43-54.




Ukrainian, with English extended summary, titles of tables, captures to figs, and references


The territory of the Skole Beskyds National Park is one of the most densely forested regions of the Ukrainian Carpathians, little affected by human activity and having a high landscape diversity and a considerably rich fauna. Investigation of the alien component of the local fauna and analysis of its impact on indigenous species, habitats and ecosystems is an urgent problem that requires discussion. The industrial activity within the Skole Beskyds was accompanied by a constant increase in anthropogenic impact on forest ecosystems of the park. Changes in the natural coenotic structure of forests led to a significant contraction of ranges of several species of the local mammal fauna and the areas became accessible for other species. The consequence of the appearance of alien species in a completely new area are the changes in composition of zonal complexes due to the loss of the rare components and the gradual transition of the zonal complexes to the gray biota state. The article analyzes the occurrence of alien species that became possible under anthropogenic conditions and depended on humans (programs for the enrichment of hunting grounds, experiments on the introduction of new species, etc.), as well as those caused by spontaneous spread and invasion. There are five groups of adventive species in the composition of the local fauna expansive (Eptesicus serotinus, Martes foina, Sciurus vulgaris carpathicus); re-appeared (Capreolus capreolus, Lutra lutra, Meles meles, Sus scrofa); invasive (Mus musculus, Rattus norvegicus); reintroduced (Bison bonasus, (Rupicapra rupicapra), introduced (Oryctolagus cuniculus, Myocastor coypus, Ondatra zibethicus, Cervus nippon, Nyctereutes procyonoides, Dama dama). Projects on the enrichment of the territory's fauna with alien species ended in failure. Thus, under the parks conditions, indigenous species show a high ability to restore population abundance. Introduction of only three species in the territory of the park was successful raccoon dog (Nyctereutes procyonoides) from the family Caniformes, muskrat (Ondatra zibethicus)from the family Muriformes and fallow deer (Dama dama) from the family Cervidae.


mammals, alien species, species diversity, protected ecosystems, Skolivski Beskydy, Ukrainian Carpathians.



Barkasi, Z. 2016. Endemism in the Mammalian fauna of the Carpathians. Proceedings of the Theriological School, 14: 315.
Barkaszi, Z. 2018. Changes in the rodent fauna (Mammalia, Glires) of the region of the Ukrainian Carpathians during the XIXXXI centuries. Geo &Bio, 16: 4862.
Bilokon, S., M. Belokon, Yu. Belokon, I. Dykyy. 2014. Genetic diversity of red squirrel (Sciurus vulgaris L.) from Western Ukraine by ssr loci. Visnyk of the Lviv University. Series Biology, 65: 296305. (In Ukrainian)
Chudiiovych, I. 2010. To the History of Hunting and Fishing in the Skolivshchyna, 1600. (web-site Hunting of Ukraine: (In Ukrainian)
Dejneka, A. M., V. Ya. Banderych, A-T. V. Bashta [et al.]. 2008. National Park Skole Beskids: Fauna. SPOLOM, Lviv, 1176. (In Ukrainian)
Didukh., Ya. 2007. Energy problems of ecosystems and provision of steady development of Ukraine. Visnyk of NAS of Ukraine, 4: 312. (In Ukrainian)
Duffey, E. 1973. Ochrona przyrody ozywionej w rezerwatach Wielkiej Brytanii. Ochrona Przyrody, 38: 2739.
Dykyy I.V., . Martsiv, V. Shelvinskiy, A. Zatushevsky. 2017. The feeding peculiarities of the Mustelidae family on the territory of the Lviv region. The Journal of V. N. Karazin Kharkiv National University. Series Biology, 29: 135141. (In Ukrainian)
Dykyy, I. V. 2004. Badger (Meles meles L., 1758) in Western Ukraine: Morphology, Distribution, Ecology and Protection. Abstract Diss. ... Cand. Biol. Sci. (03.00.08). Schmalhausen Institute of Zoology, NAS of Ukraine, Kyiv, 120. (In Ukrainian)
Gerenchuk, K. I. (ed.). 1968. The Nature of the Ukrainian Carpathians. Publishing House of Lviv University, Lviv, 1267. (In Ukrainian)
Glowacinski, Z. 2011. Karpackie kregowce Vertebrata w swietle kryteriow Czerwonej Ksiegi. Roczniki Bieszczadzkie, 19: 181190.
Holubets, M. A. 2000. Ecosystemology. Polly, Lviv, 1316. (In Ukrainian)
Ivashkiv, I. M., A.-T. V. Bashta. 2011. Specific composition and features of the spatial distributing of bats in the urboecosystems of Beskids. Scientific Bulletin of National Forestry University of Ukraine, 21 (16): 123126. (In Ukrainian)
Kazimir, . ., P. B. Khojetsky. 2004. Occupation of Caprolus caprolus L. in lands Lviv region. Scientific Bulletin of National Forestry University of Ukraine, 14 (4): 4649. (In Ukrainian)
Khojetsky, P. 2003. State of the Bison bonasus L. population in Skolivski Beskydy. Visnyk of Lviv University. Biology Series. 32: 128133. (In Ukrainian)
Khoyetskyy, P. 2012. Forest and hunting management in Western Ukraine. Abstract Dis. ... Doct. Agricult. Sci. 06.03.03. National Forestry University of Ukraine, Lviv, 40. (In Ukrainian)
Makushenko, M. O., I. D. Shnarevych. 1954. To the distribution and ecology of some types of industrial animals of the Chernivtsi region. Scientific notes of the Lviv Scientific Natural History Museum of the Academy of Sciences of the USSR, 3: 7789. (In Ukrainian)
Nowak, E. 1971. O rozprzestrzenianiu sie zwierzat i jego przyczynach. Zeszyty Naukowe Instytutu Ekologii PAN, Nr 3: 1255.
Pimentel, D. (ed.) 2002. Biological Invasions; Economics and Environmental Costs of Alien Plant, Animal, and Microbe Species. CRC Press, Boca Raton, London, New York, Washington DC, 1369.
Pimentel, D., R. Zuniga, D. Morrison. 2004. Update on the environmental and economic costs associated with alien-invasive species in the United States. Ecological Economics, 52: 273288.
Protsiv, O. R., P. B. Khoietskyi. 2010. Serna (Rupicapra rupicapra Linnaeus, 1758). Lviv, 156. (In Ukrainian)
Pysek, P., P. E. Hulme, W. Nentwig. 2009. Glossary of the Main Technical Terms Used in the Handbook. In: Handbook of Alien Species in Europe. Springer, Netherlands, 397379.
Rizun, E., V. Bondarenko. 2016. Trends in population dynamics of the Ukrainian game mammal fauna and propositions on improvement of its census. Proceedings of the Theriological School, 14 (Mammal Ecology and Geography): 3440. (In Ukrainian)
Sharapa, O. S. P. B. Khoietskyi, V. M. Smagol. 2010. Prospects for the restoration of bison (Bison bonasus L.) in the National Park Skolivsky Beskydy. Scientific Bulletin of National University of Life and Environmental Scienes of Ukraine, (147): 353361. (In Ukrainian)
Sokur, I. T. 1961. Historical Changes and the Use of Mammals in Ukraine. Ukr. Acad. Sci. Press, Kyiv, 184. (In Ukrainian)
Stetsula, N. . 2014. Biotop preferences of mouse-like rodents on the territory of the National Park Skolivski Beskydy. Visnyk of Zaporizhzhya National University. Biological Sciences. 1: 99106. (In Ukrainian)
Stoiko S., E. Hadach, T. Simon, S. Mikhalik. 1991. Protected ecosystems in the Carpathians. Svit, Lviv, 1247. (In Ukrainian)
Stoyko S.M. 2008. Biological diversity preservation, optimisation of ecological security and sustainable development in the Carpathians. Scientific bulletin of Uzhhorod University. Series Biology, 24: 510. (In Ukrainian)
Tatarynov, K. A. 1956. The Vertebrate Fauna of Ukraine. Academy of Sciences of the USSR, Kyiv, 1188. (In Ukrainian)
Tatarynov, K. A. 1973. The vertebrate fauna of Ukraine. Lviv University, Lviv, 1126. (In Ukrainian)
Urban, P., M. Uhrin. 2014. Draft Carpathian Red List of threatened Mammals (Mammalia). Carpathian Red List of Forest Habitats and Species, Carpathian List of Invasive Alien Species (draft). The State Nature Conservancy of the Slovak Republic, Banska Bystrica, 221227.
Volokh, A. M., N. V. Rozhenko. 2009. Eurasian otter. Lutra lutra (Linnaeus, 1758). In: Akimov, I. A. (ed.). Red Data Book of Ukraine. Animals. Global Consulting, Kyiv, 485. (In Ukrainian)
Zagorodniuk, I. 2001. Composition and historical changes of carnivore fauna of Ukraine. In: Large carnivores in Ukraine and adjacent countries. Ed. by I. Zagorodniuk.Kyiv, 1417. (Series: Novitates Theriologicae, Pars 4). (In Ukrainian)
Zagorodniuk, I. V. 2004. Mountain regions as zones with the highest species diversity of terrestrial vertebrates in Ukraine. Scientificnotes of Taurida National V. I. Vernadsky University. Series Biology and Chemistry, 17 (56), 2: 3338. (In Ukrainian)
Zagorodniuk, I. 2006. Adventive mammal fauna of Ukraine and a significance of invasions in historical changes of fauna and communities. Proceedings of the Theriological School, 8 (Fauna in Anthropogenic Environments), 1847. (In Ukrainian)
Zagorodniuk, I. V. 2009. Taxonomy and nomenclature of the non-Muroidea rodents of Ukraine. Proceedings of Zoological Museum, 40: 147185. (In Ukrainian)
Zagorodniuk, I. V., I. G. Emelyanov. 2012. Taxonomy and nomenclature of mammals of Ukraine. Proceedings of the National Museum of Natural History, 10: 530. (In Ukrainian)
Zawidzka, E. 1958. Geographical distribution of the dark phase of the squirrel (Sciurus vulgaris fuscoater Altum) in Poland. Acta Theriologica, 2 (8): 160174.
Zizda Yu. 2008. Changeability of colouring of fur and analysis of distribution of different subspecies of Sciurus vulgaris. Scientific Herald of Uzhhorod National University. Series Biology, 22: 212218. (In Ukrainian)
Zizda, Yu. 2006. Estimation of a diversity of color forms of the red squirrel (Sciurus vulgaris) in synanthropic and natural habitats in the Transcarpathian region. Proceedings of the Theriological School, 8 (Fauna in Anthropogenic Environments), 126132. (In Ukrainian)
Zizda, Yu. 2005. Distribution of the colored forms of squirrel (Sciurus vulgaris) in the Transcarpathians and adjacent regions of Ukraine. Scientific Herald of Uzhhorod National University. Series Biology, 17: 147154 (In Ukrainian)


to main page of journal >>>

created: 16.06.2019
updated: 01.10.2019

Locations of visitors to this page